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Manuel, M. Barnett, R. Sandoval-Velasco, M. Yamaguchi, N. Vieira, F. G. Mendoza, M. L. Z. Liu, S. Martin, M. D. Sinding, M-S. Manuel, M. d. Ross, B. Sandoval-Velasco, M. Yamaguchi, N. Vieira, F. G. Mendoza, M. L. Z. Liu, S. Martin, M. D. Sinding, M.-H. Bertola, L. D. Jongbloed, H. Van Der Gaag, K. J. De Knijff, P. Yamaguchi, N. Hooghiemstra, H. Bauer, H. Henschel, P. White, P. A. Driscoll, C. A. Tende, T. Ottosson, U. Saidu, Y. Vrieling, K. de Iongh, H. H. (2016). "Phylogeographic designs in Africa and High Resolution Delineation of genetic clades in the Lion (Panthera leo)". Kitchener, A. C. Breitenmoser-Würsten, C. Eizirik, E. Gentry, A. Werdelin, L. Wilting, A. Yamaguchi, N. Abramov, A. V. Christiansen, P. Driscoll, C. Duckworth, J. W. Johnson, W. Luo, S.-J. Meijaard, E. O'Donoghue, P. Sanderson, J. Seymour, K. Bruford, M. Groves, C. Hoffmann, M. Nowell, K. Timmons, Z. & Tobe, S. (2017). "A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group" (PDF). Burger, J. Rosendahl, W. Loreille, O. Hemmer, H. Eriksson, T. Götherström, A. Hiller, J. Collins, M. J. Wess, T. & Alt, K. W. (2004). "Molecular phylogeny of the extinct cave lion Panthera leo spelaea" (PDF)

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